According to this article...
https://expeditions.fieldmuseum.org/...nd-environment
"Ants first appeared on the earth between 140 to 168 million years ago, during the Jurassic period"
Most insects produce eggs continuously so defining the number of generations they've experienced to date is difficult. However, at even a seemingly low rate of, say, ten generations per year, that's a billion or so generations - plenty of time to fine tune some incredible behaviors.
Also, ants communicate with each other. Who's to say there aren't "captain" ants that direct worker ants to perform certain cooperative actions?
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